Hello,
You can get linux to do all that you are asking for: print server, mail server... but the integration of Linux into your environment will take a bit of experience, and I would not recommend it on your own, unless you have time to invest on learning it yourself. I would be calling in a consultant to take care of it for you, and can teach you how to do it successfully.
You can setup linux to work with Netware. Not extremely robust, but it does work. You can also have linux use Samba so that your WIndows computers can work with the fileshares too.
You will need 2 network cards if you want to work with an internal network, and an external network, and assuming that you want the linux box to also be your firewall. Do not forget to setup IPTABLES to properly firewall the internet connection.
For internet connections, you will need to use either SSH or VNC. SSH is encrypted, VNC is not. Both work really nicely. SSH for command line management and VNC for graphical management.
Don't forget to design in your backup solutions.
Christian
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