Hrm... Where to begin...
First of all, can the machines ping each other? Sometimes they can see one another through NETBIOS, which is a seperate protocol from TCP/IP. Can the Desktop ping anything on the Internet? If yes, can the laptop ping the IP address of the desktop?
Pretty much, you'd troubleshoot this like you would any router. See if the router can ping an Internet site, see if the client can ping the Internet-facing interface on the router. That should help you determine in some way or another what's going on. It may boil down that you need to uninstall/reinstall ICS on the desktop system.
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